2018–2023. Head of the Department of Evolution, Ecology & Behaviour Professor of Zoology, University of Liverpool 2013. C.A. Wright Medal, Brisitsh Society for Parasitology 1998–2018. Professor of Zoology (2004), Reader (2003), Senior Lecturer (2002), Lecturer School of Biological Sciences, University of Bristol 2011–2012. Fellow Wissenschaftskolleg zu Berlin (Institute for Advanced Study) Leader of Adaptive Plasticity working group 1997 Visiting Scientist, California Institute of Technology, USA 1989 – 1998. Research Fellow, Senior Research Fellow & MRC Career Development Award, Division of Biological Sciences, University of Edinburgh 1985 – 1989. Research Assistant, Liverpool School of Tropical Medicine 1985-1989. PhD, Liverpool School of Tropcial Medicine